Paleobiologist Emma Russell works for crypto-zoological organization Monarch to track down and study Titans, giant monsters that once dominated the Earth. She and her daughter Madison witness the birth of a giant larva dubbed Mothra. Emma calms Mothra down with a device, the “Orca”, which is capable of emitting frequencies that only Titans can hear. Mothra becomes docile, until an organization of eco-terrorists, led by Alan Jonah, attack and kidnap both Emma and Madison. Mothra manages to escape and cocoon herself underneath a waterfall.
Monarch scientists and soldiers led by doctors Ishirō Serizawa and Vivienne Graham approach Mark, Emma’s ex-husband, to track down Emma and Madison. They follow Godzilla to Antarctica and realize Jonah intends to free a Titan encased in the ice called “Monster Zero”. Jonah’s team ambushes the Monarch soldiers and Mark is unable to rescue Emma and Madison. Emma frees and awakens Monster Zero. The Titan begins rampaging until Godzilla emerges and the two engage in a fight, which ends with Monster Zero killing Graham and flying away.
The new story follows the heroic efforts of the crypto-zoological agency Monarch as its members face off against a battery of good-sized monsters, including the mighty Godzilla, who collides with Mothra, Rodan, and his ultimate nemesis, the three-headed King Ghidorah. When these ancient superspecies, thought to be mere myths, rise again, they all vie for supremacy, leaving humanity’s very existence hanging in the balance.
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